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 Salticidae (Araneae) genera of the world - an atlas
(unfinished manuscript)
by Jerzy Prószyński

Professor Emeritus, Museum and Institute of Zoology, Polish Academy of Sciences,
ul. Wilcza 63, 00-679 Warsaw, POLAND

e-mail: jerzy.proszynski@wp.pl

Chapter 23
EUODENINES
informal group of genera
Version June 8th, 1010


symbol of the supragroup HISPONINES-EUOPHRYOIDA

Composition and searching on this page: Agorius Ashtabula Bagheera Bryantella, Capeta Cerionesta Chirothecia Coccorchestes Consingis Gambaquezonia Gastromicans Itata Macaroeris Massagris Mikrus Semorina Omoedus Synagelides .

Exemplary representatives of the group
Figure - . Macaroeris nidicolens: male and female. SOURCES: A-C - Proszynski. Ann. zool. 2003: 86-87, figs 330-337) M2-M3 - same © Photo A. Senglet.: M4-M5 - photos by © J. Lissner. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

EUODENINES informal group of genera
Database contains 16 recognizable genera,139 species
name combines of parts of names of related groups EUO-phryines-DEN-dryphant-INES].

This informal group was assembled for logistic convenience, to facilitate handling of too numerous genera of EUOPHRYINES. It contains diverse genera departing from average EUOPHRYINES and DENDRYPHANTINES, is pending for further research. Some males have enormous single coil of embolus, often shifted to the lateral surface of bulbus (like in Bagheera Fig. C), or have atypical spermophor, straight or gently waving (Capeta Fig. A), or have otherwise incompatible palps (like in Synagelides Fig. D, or Agorius Figs N-P). Palp of Macaroeris Fig. A has typical meandering spermophor of EUOPHRYINES, but its epigyne (Fig. F) fit rather DENDRYPHANTINES. Ducts and spermatheca of Gambaquezonia are most unusual (Figs C, G-H).

Constelation of supposed
Gen. Agorius Thorell, 1877
Type species Agorius gracilipes (11 recognizable species)
See more species at
Agorius-Q+M

Remarks. Agorius, like many other unrelated genera of Salticidae, has elongated, constricted abdomen and particularly long leg I, with characteristic group of spines ventrally on metatarsus I (Fig. P). Differences in palps and epigyne are so big that particular species are apparently not congeneric and may represent three, or more unrelated genera. Type species is apparently not congeneric with other "Agorius" species, its placement could not be determined without description of matching male.

A - Agorius gracilipes, B-D - Agorius lindu, E-G - Agorius tortilis, H-I - Agorius cf. borneensis, Sabah: Trus Madi, 846, J-K - Agorius sp. F. Sabah: Danum Valley, 903, L - Agorius borneensis, M - Agorius kerinci, N, R - Agorius constrictus, O-Q - Agorius saaristoi, S-U - diversity of Agorius sp. (S-T - from Singapore, U - from Halmahera: Tanah Putih NP).
SOURCES: A - Proszynski 1968b. Ann. zool., 26: 221-225, ff. 10-12, - Prószynski INTERNET + 2008b. Adv. Arachn.... Inst. Zool., bulg. Acad. Sci. Monogr. 12: 321-322, f 7, 8, 29, 30, 41, 54, 59, E-G -Cao & Li, sp. n. ZooKeys 630: 46-51, f. 3–4, 43, H-I - Photo © P. Koomen, L - Edmunds, Prószynski. 2001. Bull. Brit. Arachn. Soc. 12 (3): 139-140. f 1-8, Prószyński, 2008b: 12: 322, Figs. 5, 20, 34, 44, S-T - ©Photo H.K. Tang, U - ©Photo D. Knowles. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Ashtabula Peckham & Peckham, 1894
Type species Ashtabula zonura (9 recognizable species)
See more species at
Ashtabula-Q+M

A26
A - Ashtabula zonura, B-C - Ashtabula dentichelis, D - Ashtabula dentata, E - Ashtabula bicristata.
SOURCES: A - Peckhams 1894. Occ. Pap. Nat. Hist. Soc. Wisc., 2(2): 140, t 14, f 4, B-C - Maddison 1996: 154 (4): 336, fig. 102, D - Chickering 1946. Bull. Mus. Comp. Zool. 97: 248-251, figs 206-211, E - Galiano 1972b. Revista Mus. argent. Entomol. 4 (1): 39-41, f. 78. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Bagheera Peckham, Peckham, 1896
Type species Bagheera kiplingi (4 recognizable species)
See more species at
Bagheera-Q+M

A-E - Bagheera kiplingi.
SOURCES: A-B - Maddison 1996.Bull.Mus.Comp. Zool. 154(4): 233, 335, 337, f 71, 80-83, C-E -©Photo by Eric Olson.. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Bryantella Chickering, 1946
Type species Bryantella speciosa. (x recognizable species)
See more species at
Bryantella-Q+M

A - Bryantella speciosa, B - Bryantella convexa, C-D - Bryantella smaragda, E - Bryantella tropica, F - Metaphidippus mandibulatus for comparison.
SOURCES: A -B - Chickering, 1946: 335, f. 292-297, C-D - Crane J. 1945. Zoologica, 30 (1, 3): 40, figs 5a-d, E -F - Maddison 1996. Bull. Mus. Comp. Zool., 154(4): 238, f. 62 ; 231, 234, 335, f 74, 96-98 All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Nomenclatorical correction
Metaphidippus tropicus = Bryantella tropica Peckham & Peckham, 1901) (compare Figs E -F).

Gen. Capeta Ruiz & Brescovit, 2005
Type species Capeta tridens .(2 recognizable species)
See more species at
Capeta-Q+M

C
A-B - Capeta tridens, C-D - Capeta cachimbo.
SOURCES: A-B - Ruiz , Brescovit. 2005a. Revta bras. Zool. 22: 692, f 21-24, C-D - Ruiz, Brescovit 2006a. Revta bras. Zool. 23(2): 353-354, f 9-12.. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Cerionesta Simon, 1901
Type species Cerionesta luteola (1 recognizable species)
See more at
Cerionesta-Q+M

C-C5 - Cerionesta luteola.
SOURCES: C - Galiano, Physis C. 1976, 35(90) : 59-61, f 1-7, C5 - Peckhams 1893: 700, f 8-e. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Chirothecia Taczanowski, 1878
Type species Chirothecia clavimana (x recognizable species)
See more species at
Chirothecia-Q+M

A-C - Chirothecia clavimana, D - Chirothecia wrzesniowskii [epigyne does not agree with type species = misplaced!].
SOURCES: A-C - Galiano 1972b. Rev. Mus. argent. Cien. natur. Entomol., 4 (1): 5-12, ff 13-16, 23-26. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Coccorchestes Thorell, 1881
Type species Coccorchestes rufipes.(x recognizable species)
See more species at
Coccorchestes-Q+M , compare also Porius.

A-C - Coccorchestes rufipes [copulatory openings blocked by waxy stopper?], D-F, K-M - Coccorchestes ferreus, G - Coccorchestes taeniatus, , H-I - Coccorchestes buszkoae, J - Coccorchestes waris. N - C. cf. clavifemur.
SOURCES: A-C -- Proszynski 1971c. Ann. zool. 28: 156-160, ff. 1-14, 15-16,K-M -- ©Photo G. Anderson, D-F - Davies Todd V., Zabka M. 1989. Mem. Queensland Mus., 27 (2): 229, 230, t. 33, G, J, - Balogh P. 1980a. Acta Zool. Acad. Sci. Hung., 26(1-3): 78, ff. 33-34, 93, N -- ©Zhang J., Maddison 2015: Zootaxa 3938 (1): f 828 +d) f 826-827. © Magnolia Press.. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Consingis Simon, 1900
Type species Consingis semicana (1 recognizable species)
See more species at
Consingis-Q

A-D - Consingis semicana.
SOURCES: A-D - Galiano 1963b. Physis, 23 (66): 331, t. 16, ff. 3, 4. + 1974a. Physis C., 33 (86): 2-5, f 24, 25. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Gambaquezonia Barrion, Litsinger, 1995
Type species Gambaquezonia itimana.(3 recognizable species)
See more species at
Gambaquezonia-Q+M

A-B - Gambaquezonia itimana (holotype specimen and internal structure of its epigyne), C-F - Gambaquezonia sp.(misidentified as G. itimana), G-L - Gambaquezonia curioi (PHILIPPINES: Panay).
SOURCES: A - Barrion, Litsinger 1995: 95-96, figs 49a-e, B - Proszynski 1995-2016, Internet Database of Salticidae - internal structure of epigyne of the holotype specimen, C-F - Edwards (2007) J. Arachnol. 37: 103-105, f 1-10, G-L - Freudenschuss, Grabolle, Krehenwinkel, 2016 Arachnology 17(1): 25-27, figs 1-8, and ©Photo M. Freudenschuss. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.
ATTENTION: Fig. B showing internal structure epigyne of the holotype specimen contributed to paper of Edwards (2007) by Prószyński, unfortunately Dr. Edwards did not understand its significance and failed to publish it.

Gen. Gastromicans Mello-Leitão, 1917
Type species Gastromicans albopilosa (x recognizable species)
See more species at
Gastromicans-Q+M

A - Gastromicans albopilosa, B-C - Gastromicans levispina, D - Gastromicans sp. [?], E - Gastromicans sp. [?].
SOURCES: A - Galiano 1963b. Physis, 23 (66): 306-307, t 12, f 1-2, B-C - Maddison W.P. 1996. Bull.Mus. Comp. Zool., 154 (4): 232, 237, 335, f. 73, 93-95, 101 ©Magnolia Press, D - Maddison 2015. Journal of Arachnology. 43: 231–292, f. 75, E - ©Photo Gasnier & Machado. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Itata Peckham, Peckham, 1894
Type species Itata vadia (3 recognizable species)
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Itata-Q+M

A - Itata vadia, B- Itata tipuloides, C-D - Itata sp.
SOURCES: A - Peckhams 1894.Occ. Pap. Nat. Hist. Soc. Wisc., 2 (2): 115, t 11, f 5, B - Galiano M.E. 1963b. Physis, 23 (66):: 378-379, t. 25, ff. 1-3, C-D - ©Photo Gasnier & Azevedo. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Macaroeris Wunderlich, 1992
Type species Macaroeris nidicolens (x recognizable species)
See more species at
Macaroeris-Q+M

A-D - Macaroeris nidicolens from Israel (C-D - teratological distorted epigyne or different species), D-E - epigyne specimen from Europe.
SOURCES: A-D - Proszynski. Ann. zool. 2003: 86-87, figs 330-337, M2-M3 - © Photo A. Senglet.
Diversity in Macaroeris - males and females: A-D - Macaroeris diligens, E-F: - Macaroeris moebi (Photos of - Macaroeris nidicolens, male and female - see top of this chapter, Figs D and E).
SOURCES: All © photos by J. Lissner. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Massagris Simon, 1900
Type species Massagris constricta (7 recognizable species)
See more species at
Massagris-Q+M

Remarks. Type species of the genus - Massagris constricta is considered lost, D. J. Clark (1974: 20) was most probably the last person who has seen it - according to Wesołowska (1993: 140). However, some specimens borrowed by Clark were returned to owners after his death (see introduction to Clark 1974: 12), so one cannot exclude possibility that specimens of M. constricta (Simon's coll. No. 9925) was mixed up and left in the spider collection of Natural History Museum in London, or else was returned to Museum National d'Histoire Naturelle in Paris, but was placed there in a wrong jar, so it was not recovered later. I would advice persons checking Salticidae collections in both Musei to remember on that possibility.

A - Massagris constricta, B-C - Massagris honesta, D - M. natalensis, E - M. honesta.
A - Massagris constricta, B-C - Massagris honesta, D - M. natalensis, E - M. honesta, F-H - Massagris mohale from Lesotho, I- "Massagris" schisma [misplaced!].
SOURCES: A - Simon 190a. 451, 452, B-C - Wesołowska & Haddad 2013. Afr. Invert.: 54 (1): 210-212, f 19, 90, 91, 107–113, D-E - Photo Hadad from Wesołowska & Haddad 2013, F-H - Wesolowska W., Haddad C.R., 2014. Afr. Invert. 55 (2), 250-253, f 69, 70, 74–81, I - Maddison & Zhang, 2006 :Zootaxa 1255: 32, f. 8-10 Magnolia Press.. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Mikrus Wesołowska, 2001
Type species Mikrus ugandensis (1 recognizable species)
See more species at
Mikrus-Q+M

Mikrus ugandensis. ATTENTION: type of palp resembles Holoplatys.
SOURCES: Wesolowska W. 2001c. Genus, Wroclaw, 12(4): 585-588, figs 1-7. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Omoedus Thorell, 1881
Type species Omoedus niger (4 recognizable species)
See more species at Omoedus-Q+M

REMARKS. Transfer from EUOPHRYINES tentative, based on difference in structure of spermathecae and ducts, while palp is typical for EUOPHRYINES and BALLINES. Recognizable by internal structure of epigyne, single males, and females with different epigyne cannot be identified as Omoedus, part of references in the WSC is uncertain and require revision.

Interpretation of alleged synonymy of genera Omoedus with Pystira and Zenodorus by Zhang & Maddison, 2012b: 21 caused deep clevage between students of Salticidae because of consequences to methodological definition of a genus. See diagnostic characters of Omoedus and Zenodorus, below.

A-B - Omoedus niger type specimen, C - Omoedus sp. D, F - Omoedus[?] insultans, H-I - Omoedus piceus & Omoedus cf. piceus.
SOURCES: A-B, E, H-I - Proszynski J. 1971c. Ann. zool. 28: 172-175, f 28-40, C - © Photo D. Knowles, F-G - Hęciak, Prószynski 1984b: 411, f 2, 5, 10, D - ©Zhang J., Maddison 2015: Zootaxa 3938 (1): 26, f 800. © Magnolia Press. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Peckhamia Simon, 1901
Type species Peckhamia scorpionia.(6 recognizable species)
See more species at
Peckhamia-Q+M

REMARKS. Placement uncertain due to poor genitalic documentation, forked tibial apophysis of the type species (with absence of other details of palp and epigyne) suggest placement rather in COLONINES, other species are most probably EUODENINES.

A - Peckhamia scorpionia, B-D - Peckhamia picata, E - Peckhamia sp.
SOURCES: A - Peckhams 1909. 16 (1): 370, t 50, 51, f 3, B, D - Paquin, Duperre. 2003. Fabreries, Suppl. 11: 197, f 2205-2208, C - Kaston B. J. 1948. 70: 452, t 88, f 1628-1630, P4 - Maddison 2015. Journal of Arachnology. 43: 231-292, f. 68.. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Semorina Simon, 1901
Type species Semorina seminuda (4 recognizable species)
See more species at
Semorina-Q+M

A - Semorina seminuda, B - Semorina brachychelyne.
SOURCES: A - Galiano 1963b. Physis, 23 (66): 445-447, t 37, f 6-7, B - Crane 1949. Zoologica, 34 (2, 7): 35, f 2a-g. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.
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Gen. Synagelides Strand, 1906
Type species Synagelides agoriformis (36 recognizable species)
See more species at
Synagelides-Q+M

A - Synagelides agoriformis Type specimen, B-D - Synagelides agoriformis - Russia: Far East - Primore, E-H - Synagelides brahmaputra.
SOURCES: A - Proszynski J. 1979. Ann. zool. 34: 316, f 307- 315, B-D - Proszynski 1979. Ann. zool. 34: 318, f 307- 315, E-H - Caleb et al. 2018: Acta arachnologica 67(1): 9-12, f. 12-24. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.