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Proszynski's Salticidae (Araneae)
Genera of the World
vol. II - species attachments

Chapter 79

"THIRATOSCIRTINES"
species attachments

Correlated chapter in vol. I
Genera comparable with THIRATOSCIRTINES
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symbol of the genera not classified into supragroups
Contents of this page and facility for searching
Type genus: Thiratoscirtus.
Ajaraneola Alfenus Ansienulina Bacelarella "Bacelarella-2" "Bacelarella-3" Gramenca Lamottella Longarenus Malloneta Nimbarus Pochyta Saraina Tarne Thiratoscirtus Conclusion on group "Thiratoscirtines"

Introduction

Bodner & Maddison (2012, p. 221-222) proposed a new clade "THIRATOSCIRTINAE", which they consider subfamily, and discuss its evolutionary significance. Understanding premises for that is difficult because their taxonomic documentation is limited to listing genera they propose to include into THIRATOSCIRTINAE, without taxonomic definition or description, apart from a cladogram based on molecular analyses, whose taxonomic significance is unclear. To provide some orientation in their proposal I quote below the taxonomic documentation of all species they consider.

Remarks of relationships between genera of Thiratoscirtinae and other clades of Salticidae. Morphological documentation available in the literature is insufficient and therefore inconclusive. Structure of epigyne and palps represent two types, which deserve further attention, there are also some rather incompatible species. The most tempting similarity is separation of eyes anterior lateral and posterior lateral by prominent depression, with microscopic size of eyes II, supposedly indicating relationships to fossil Eocene Prolinus, and its extant analogues Tomocyrba and Hispo, looks probable in the case of Massagiris schisma, (whose relationship to Thiratoscirtinae is, however, questionable), but is not supported by eyes of Thiratoscirtus and related genera. Coiled embolus may indicate relationships to Euophryinae, but require further documentation of its basis. There is intriguing resemblance in posterior part of epigyne with Asiatic genus Habrocestoides, with curious circular structure inside, but that require further study. In view of very incomplete morphological documentation of diversity in Thiratoscirtinae, no firm conclusion on their placement could be drawn.
Good illustration of Bodner and Maddison views is the plate of 24 unidentified species - since I have no current copyright permission to display the plate, I direct readers to the original publication, including here thumbnails advertisement for that (below).
In absence of diagnosis in words - examination of genitalic characters characters permit to interpret authors' concept of Thiratoscirtines (1-3 vertical rows) in comparison with their Plexippines (the fourth vertical row). Unidentified specimens from Gabon. SOURCE: Bodner, M. R. & Maddison, W. P. (2012). The biogeography and age of salticid spider radiations (Araneae: Salticidae). Molecular Phylogenetics and Evolution 65(1): 213-240.Bodner, Maddison 2012: 65: 237-238, plate B1-2 with 24 unidentified Thiratoscirtinae [?].
Continuation - more unidentified Thiratoscirtines from Gabon. SOURCE: Bodner, M. R. & Maddison, W. P. (2012) - as above.
Remarks on genus Thiratoscirtus and related forms was given by Wesołowska 2011 (below). The documentation of Thiratoscirtus patagonicus, the type species of the genus Thiratoscirtus, prepared by Szuts and documenting its African relationships, has been reproduced in the database of Salticidae by Prószynski, 2003b (by kind permission of T. Szuts).
Definition by Wesolowska, Russel-Smith 2011. Ann. zool. 61(3): 603-604, f 188-192, 252.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Pictorial guide to identification of genera


Comparison of selected characters


Gen. Ajaraneola Wesołowska, Russel-Smith, 2011
Type species Ajaraneola mastigophora(1 species).
Compare with other genera in vol. I at Ajaraneola
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Ajaraneola mastigophora : Wesolowska, Russel-Smith.Ann. zool. 61(3): 558-559, f 14-20. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Alfenus Simon, 1902
Type speciesAlfenus calamistratus (2 species).
Compare with other genera in vol. I at Alfenus
Alfenus calamistratus : Szuts T., Scharff N. 2005. Acta zool. Acad. Sci. Hungar. 51: 359-361, f 1a,c, 2a-f, 3a-d.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.
Alfenus chrysophaeus : Szuts T., Scharff N. 2005. Acta zool. Acad. Sci. Hungar. 51: 361, f 1a,c, 2a-f, 3a-d..All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Ansienulina Wesołowska, 2015
Type species Ansienulina mirabilis (1 species).
Compare with other genera in vol. I at Ansienulina
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Ansienulina mirabilis: Wesolowska, 2015: 56(2): 478, f. 1-14.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Bacelarella Berland, Millot, 1941
Type species Bacelarella fradei (4 species)
Compare with other genera in vol. I at Bacelarella
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Bacelarella fradei : Proszynski 1987. Atlas: 4. [Syntypes] + Rollard, Wesolowska. 2002. Zoosystema. 24 (2): 286, figs 2A-E + Szuts, Jocque 2001. Annls Mus. R. Afr. Centr. (Zool). 285: 88-91, figs 3F, 3M, 5B, 5E, 7B, 7E, 8C, 8F, 9D. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.
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Bacelarella dracula: Szuts, Jocque 2001. Annls Mus. R. Afr. Centr. (Zool). 285: 86-88, figs 1B, 2A, 3D, 3K, 4C, 4D, 4G, 4H, 7C, 7F, 8B, 9B + Wesołowska, Russel-Smith 2011. Ann. zool. 61(3): 559-561, f 217-218. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.
Bacelarella iactans : Szuts, Jocque 2001. Annls Mus. R. Afr. Centr. (Zool). 285: 90-91, figs 3E, 3L,5A, 5D,6, 8C, 9C..
Bacelarella tanohi : Szuts, Jocque 2001. Annls Mus. R. Afr. Centr. (Zool). 285: 91-92, figs 1A, 3G, 3N, 5C, 5F, 8E..All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Bacelarella-2 Berland, Millot, 1941
Type species Bacelarella conjugans (4 species)
Compare with other genera in vol. I at Bacelarella
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"Bacelarella" conjugans : Szuts, Jocque 2001. Annls Mus. R. Afr. Centr. (Zool). 285: 83-85, figs 1C-D, 2B-E, 3C, J, 4B, F, 7A, D, 8A, 9A. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.
b)
"Bacelarella" pavida +b) tentativa: Szuts, Jocque 2001. Annls Mus. R. Afr. Centr. (Zool). 285: 83-85, figs 3B, 3I, 4A, 4E +b) 285: 82-83, figs 3A, H .All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.
"Bacelarella-2" gibbosa : [third genus?] Wesołowska, Edwards 2012. Ann. zool.: 62(4): 735, f 5–11, 111–113.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. "Bacelarella-3"[?]
Representative species "Bacelarella"gibbosa (4 species)
Compare with other genera in vol. I at Bacelarella-3
different genus, to be described as new.
Plate XXX. 756. A-D - "Bacelarella-3" gibbosa : [third genus?].
SOURCE. Wesołowska, Edwards 2012. Ann. zool.: 62(4): 735, f 5–11, 111–113. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Gramenca Rollard, Wesołowska, 2002
Type species Gramenca prima (1 species)
Compare with other genera in vol. I at Gramenca
Gramenca prima : Rollard Ch., Wesolowska W. 2002. Zoosystema. Paris, 24 (2): 292-293, figs 7A-D.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Lamottella Rollard, Wesołowska, 2002
Type species Lamottella longipes (1 species);
Compare with other genera in vol. I at Lamottella
Lamottella longipes : Rollard Ch., Wesolowska W. 2002. Zoosystema. 24 (2): 296, figs 10A-F.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Longarenus Simon, 1903
Type species Longarenus brachycephalus (1 species)
Compare with other genera in vol. I at Longarenus
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Longarenus brachycephalus + Longarenus sp.:: Szuts T. 2005 [2007]. Opusc. zool., 36: 89-90, f 16-21 + Wesolowska, Edwards 2012. Ann. zool.: 62(4): 748, f 51–59, 124. Maddison 2015. Journal of Arachnology. 43: 231–292, f. 122.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy..

Gen. Malloneta Peckham, Peckham, 1901
Type species Malloneta guineensis (1 species)
Compare with other genera in vol. I at Malloneta
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Malloneta guineensis : Proszynski J. 1987. Atlas ...: 62. [Lectotype, Freetown [Sierra Leone] + Wesolowska, Edwards 2012. Ann. zool.: 62(4): 750-751, f 60–62, 125.Malloneta sp.: Maddison 2015. Journal of Arachnology. 43: 231–292, f. 124.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy

Gen. Nimbarus Rollard, Wesołowska, 2002
Type species Nimbarus pratensis (2 species).
Compare with other genera in vol. I at Nimbarus
Nimbarus pratensis : Rollard Ch., Wesolowska W. 2002. Zoosystema. Paris, 24 (2): 301-303, f 14A-F.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Comparison of palps of 5 species, used by Maddison et al., 2008 for gene sequencing (upper row), compared with type specimes with types of genera they supposedly belong (bottom row). Original captions enclosed, could be read after magnification of screen, by even 400%.

Gen. Pochyta Simon, 1901
Type species Pochyta spinosa (9 species)
Compare with other genera in vol. I at Pochyta
b)c)d)e)
Pochyta spinosa +b) simoni +c) poissonii +d) perezi +e) remyi : Proszynski J. 1987. Atlas ...: 79 [Not a type. Is it correctly identified? ] +b) Lessert R. 1925a. Rev. suisse zool., 31 (13): 505, f 87-91 +c) Berland, Millot 1941. 12, 2: 363, f 64 +d) 362 f 63 [note embolus similarity to Massagris schisma!] +e) 364, f 64. .All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.
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Pochyta pulchra : Proszynski 1984c. Atlas ...: 70. ©Photo T. Szuts.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.
d)b)c)
Pochyta fastibilis +b) major +c) moschensis +d) ["pannosa" & 6 other MISIDENTIFIED] [true"pannosa" is "species inquirenda"] : Simon E. 1903a.: 722, f 867 A-B +b) 716, f. 863-864 (=K-L) +c) Caporiacco 1947d. 40: 249, t 2, f 76 +d) Maddison, Bodner, Needham 2008. Zootaxa 1893: 52, f 5. ©Magnolia Press. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Ragatinus Dawidowicz & Wesołowska, 2016
Type species Ragatinus maddisoni Dawidowicz & Wesołowska, 2016: 455, f. 71-78, 108-110 (1 species)
Compare with other genera in vol. I at Ragatinus
Plate XXX. 762. A - Ragatinus maddisoni : 455, f. 71-78, 108-110.SOURCE. Dawidowicz, A. & Wesołowska, W. (2016). Jumping spiders (Araneae: Salticidae) of Kenya collected by Åke Holm. Annales Zoologici, Warszawa 66(3): 437-466. . All ©copyrights are retained by the original authors and copyright holders, used by their courtesy .ragatinus_maddisoni

Gen. Saraina Wanless & Clark, 1975
Type species araina rubrofasciata (3 species)
Compare with other genera in vol. I at Saraina
Saraina rubrofasciata : Szuts T., Scharff N. 2005. Acta zool. Acad. Sci. Hungar. 51: 366-368, f 5a-g, 6a-c + Azarkina 2009: 293, FIG. 1.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.
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Saraina deltshevi +b) kindamba: Azarkina 2009: 293-295, figs 2-8 +b) 295-296, figs 9-18.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Tarne Peckham, Peckham, 1901
Type species Tarne dives(1 species)
Compare with other genera in vol. I at Tarne
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Tarne dives : Proszynski 1987. Atlas ...: 106. + Wesolowska, Edwards 2012. Ann. zool.: 62(4): 759, f 88–94, 129 + Szuts, Rolard 2007. Insect Syst. Evol, 38: 427-432, t 1a-g.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Plate XXX. 738. A-E - Tauala lepidus, F-I - Tauala sp.SOURCE. A-C - Davies Todd V., Zabka M. 1989. Mem. Queensland Mus., Brisbane, 27 (2): 209, t 15, D-I - ©Photo by R. Whyte, G. Anderson. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.

Gen. Thiratoscirtus Simon, 1886
Type species Thiratoscirtus patagonicus (17 species)
Compare with other genera in vol. I at Thiratoscirtus
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Thiratoscirtus patagonicus : Galiano 1963b. Physis, 23 (66): 457-458, t. 39, f 11-14 + ©Photo Szuts [type specimen + its label + new African specimens].All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.
b)<>c)
Thiratoscirtus rudyi [undescribed] +b) procerus + c) Thiratoscirtus [?] sp. : ©Photo Szuts [from Congo (Brazzaville): Sibiti] +b) Wesołowska, Edwards 2012. Ann. zool.: 62(4): 761-762, f 100–101, 130. Maddison 2015. Journal of Arachnology. 43: 231&#8211;292, f. 123.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.
b)c) d)
Thiratoscirtus atakpa +b) efik [misplaced] +c) torquatus +d) alveolus : Wesołowska, Edwards 2012. Ann. zool.: 62(4): 759-760, f 95–96 +b) 760, f 97–99 +c) 762-763, f 102–107, 131
+ d)Wesołowska Russel-Smith, 2011. Ann. zool. 61(3): 599-600, f 167-169.
b)
Thiratoscirtus bipaniculus +b) mirabilis : Wesołowska Russel-Smith 2011. Ann. zool. 61(3): 600, f 170-172, 236 +b) 603-604, f 188-192, 252.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.
b)
Thiratoscirtus gambari +b) monstrum : Wesołowska Russel-Smith, 2011. Ann. zool. 61(3): 601, f 173-184, 250, 251 +b) 601-602, f 193-195, 253.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.
b)
Thiratoscirtus obudu [misplaced] +b) vilis : Wesołowska Russel-Smith 2011. Ann. zool. 61(3): 606, 196-198, 254, 255 +b) 601-602, f 199, 200, 256.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.
b)
Thiratoscirtus harpago +b) yorubanus : Wesołowska Russel-Smith 2011. Ann. zool. 61(3): 601-602, f 185-187 +b) 608-609, 201-204.All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.
Thiratoscirtus mastigophorus :Wiśniewski, Wesołowska 2013. ©Genus 24(2): 252-260, f 1-13. All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.
Thiratoscirtus perspicuus :Wiśniewski, Wesołowska 2013. ©Genus 24(2): 256-260, f 14-23..All ©copyrights are retained by the original authors and copyright holders, used by their courtesy.