RETURN TO DATABASE - ALTERNATIVE CLASSIFICATION
SALTICIDAE OF THE WORLD - 0-INDEX of Genera - 01-FOREWORD & KEY to groups of genera! - 02-Comparison - HISTORICAL classification of Aelurilleae - 03-AEURILLINES - 04-AMYCINES- 05-AMYCOIDA VARIA - 06-ASTIAINES - BALLINES - 07-BELIPPINES - 08-CHRYSILLINES - 09-COCALODINES - 10-COLONINES[=Thiodinines] - 11-DENDRYPHANTINES - 12-DIOLENINES - EUODENINES - 13-EUOPHRYINES-PART 0: Introduction-Diversity - 13-EUOPHRYINES-PART 1: Admestina-Donoessus - 14-EUOPHRYINES-PART 2: Echeclus-Pystira - 15-EUOPHRYINES-PART 3: Rhyphelia-Zenodorus - 16-EUPOAINES - 17-EVARCHINES - 18-FOSSILS - 19-HABRONATTINES - 20-HARMOCHIRINES - 21-HELIOPHANINES - 22-HISPONINES - 23-HYLLINES - 24-ICIINES - 25-LAPSIINES - 26-LIGONIPEINES - 27-LYSSOMANINES - 28-MENEMERINES - 29-MYRMARACHNINES - 30-NOTICIINES - 31-PELLENINES - 32-PSEUDICIINES - 33-SIMAETHINES - 34-SITTICINES - 35-SPARTAEINES - 35a-SYNAGELINES - 36-THIRATOSCIRTINAE - 37-YAGINUMAELLINES - 38-YLLENINES - 39-UNCLASSIFIED TEMPORARY
APPENDICES -AP1-Simon's classification - AP2-Petrunkevich synthesis-1928 - AP3-Bonnet's list of subfamilies - AP4-Prószynski's revision of subfamilies 1976 - AP5-Prószynski - summary of results - AP6-Aelurillus black & gray - AP7-Maddison's views on Salticidae phylogeny 2014 - MS1-Omoedus synonymy

MADDISON'S ALTERNATIVE: Tittle_pg Index Introduction Agoriini Aelurillina Asemoneinae Amycini Amycoida-incertae sedis Astiini Ballini Baviini Bredini Chrysillini Cocalodini Dendryphantina Euophryini-1 Euophryini-2 Euophryini-3 Eupoinae Freyina Gophoini Harmochirina Hasariini Hisponinae Holcolaetina Huriini Itatina Lapsiini Leptorchestini Lyssomaninae Marpissina Mopsini Myrmarachnini Nannenini Neonini Onomastinae Plexippina Salticini Sarindini Scopocirini Simaethina Simonellini Sitticini Spartaeina Synagelina Thiodinini Thiratoscirtina Tisanibini Viciriini Salticidae-incertae sedis

Monograph of the Salticidae (Araneae) of the World 1995-2015. Part I: Introduction to alternative classification of Salticidae.

Comparison of classificatiopns by Maddison 2015 and Prószyński 2016a
Compiled by Jerzy Prószyński
Maddison's alternative

Subfamily: Spartaeinae Tribe: Spartaeini: Subtribe: Spartaeina
M_07 - Spartaeina
Version July 1st, 2016.
Sources and permissions of illustrations are displayed in the IInd part of the monograph http://www.peckhamia.com/salticidae/

Interactive index of genera. Type genus Spartaeus.
Brettus Cocalus Cyrba Gelotia Meata Meleon Mintonia Neobrettus Paracyrba Phaeacius Portia Spartaeus Taraxella Veissela Wanlessia Yaginumanis
.

Gen. Brettus Thorell, 1895 (7 species)

Type species Brettus cingulatus.
Diagnostic illustrations below are integral part of definition.

Brettus cingulatus : Wanless 1979b. Bull. brit. Mus. nat. Hist. (Zool.): 185-187, f. 1A, C, E, G; 2A-B . By courtesy.
b)+
Brettus albolimbatus +b) cf. albolimbatus : Wanless 1979b. Bull. brit. Mus. nat. Hist. (Zool.): 188, f. 2C; 3C-D; 4A-B + ©Photo Anurag + ©Photo Vipin Baliga. By courtesy.
Brettus adonis : Wanless 1979b. Bull. brit. Mus. nat. Hist. (Zool.): 186, f. 1B, D, F, H; 2D-E; 3A.. By courtesy.
+
Brettus celebensis : Wanless 1979b. Bull. brit. Mus. nat. Hist. (Zool.): 220-221, ff. 2A-D. + Proszynski. Internet (N Sulawesi: Dumoga Bone Nat. Park. Det. Ch. Deeleman-Reinhold. By courtesy.ANALOGY: Internal structure of epigyne in Meata.
+
Brettus anchorum : Jastrzebski 1997. Genus, 8 (3-4): 702-703, figs. 1-9 + Wanless 1979b. Bull. brit. Mus. nat. Hist. (Zool.): 188, ff. 2F; 3B; 4C-D. By courtesy.
b)
Brettus madagaskarensis +b) storki : Wanless F.R. 1980a. Bull. brit. Mus. nat. Hist. (Zool.) 38 (4): 221-223, figs 3A-D +b) Logunov, Azarkina, 2007: 16(2): 98-99, f. 1-10. By courtesy.

Gen. Cocalus Koch C.L., 1846 (4 species)

Type species Cocalus concolor.
Diagnostic illustrations below are integral part of definition.

b)+
Cocalus concolor b) gibbosus : Proszynski 1984c: 158 ( Holotype) +b) Wanless 1981c. Bull. brit. Mus. nat. Hist. (Zool.), 41 (5): 256, f. 2A-F + Davies Todd, Zabka 1989. Mem. Queens. Mus., 27 (2): 196, t 4 + © ©Photo R. Whyte. By courtesy.
b)
Cocalus limbatus + b) murinus : Wanless 1981c. Bull. brit. Mus. nat. Hist. (Zool.), 41 (5): 258-260, ff. 5A-D +b) 256-258, ff. 3A-E. + © Photo H. K. Tang. By courtesy.

Gen. Cyrba Simon, 1876 (8 species)

Type species Cyrba algerina.
Diagnostic illustrations below are integral part of definition.

+
+
Cyrba algerina : Proszynski 2003a. Annales zoologici. 53 (1): 46-47, figs 153-157 + ©Photo J. Lissner (from Spain) + ©Photo B. Knoflach (from Greece) [conspecific?]. By courtesy.
+
++
Cyrba ocellata : Proszynski 1978a: 16, ff. 19, 21-24 + Davies Todd, Zabka 1989. Mem. Queens. Mus., 27 (2): 194, 198, t. 6 + ©Photo J. Caleb (Tamil Nadu, in preparation for print, ppecimen alive and discolored by preservation) + ©Photo Daiqin Li (as Paracyrba wanlessi). 2015. By courtesy.
Cyrba wanlessi [=Paracyrba w. - comb.n.] :Zabka, Kovac 1996. Senck. biol. 76(1/2): 154-156, f 1-19. By courtesy.
b)
Cyrba dotata +b) legendrei : Wanless 1984b. Bull. brit. Mus. nat. Hist. (Zool.), 47 (7): 470, ff. 15A-E +b) 458, ff. 9A-J.
+
Cyrba boveyi : Wanless 1984b. Bull. brit. Mus. nat. Hist. (Zool.), 47 (7): 463-465, ff. 11A-F + Wesołowska, Haddad 2009. African Invertebrates, 50(1): 23-26, photo 238. By courtesy.
+++
Cyrba lineata : Wesolowska 2006c. Genus 17(4): 617-620, f 1-5 + Wesołowska, Haddad 2009.Afr. Invert. 50(1): 26-27, f 24-25 +Azarkina, Logunov 2010. Afr. Invert. 50(1): . By courtesy.
++
Cyrba nigrimana : Wanless 1984b. Bull. brit. Mus. nat. Hist. (Zool.), 47 (7): 465, ff. 12A-G + Azarkina, Logunov 2010. Afr. Invert. 50(1): 167-169, f 14-23 + Haddad, Wesołowska 2009. © Afr. Invert. 50(1): 27-28, f 26-28 + Haddad, Wesołowska 2011. © Afr. Invertebr. 52(1): 59, f 5-6. By courtesy.
++
Cyrba simoni : Wesolowska W., A. Russel-Smith, 2000. Tropical Zoology, 13 (1): 20, figs 20-24 + Wanless 1984b. Bull. brit. Mus. nat. Hist. (Zool.), 47 (7): 461-463, ff. 1, 10A-L, 17E-F, 22E . By courtesy.

Gen. Gelotia Thorell, 1890 (8 species)

Type species Gelotia frenata .
Diagnostic illustrations below are integral part of definition.

Gelotia [?] sp.: Maddison 2015. Journal of Arachnology. 43: 231–292, f. 102. By courtesy. Note - body shape difference with female demands confirmation by palp documentation!
b)c)
Gelotia frenata b) bimaculata c) bouchardi : Proszynski 1968a.77: 11-16, ff. 1, 3, 5, 7, 9 +b) 16-20, ff. 2, 4, 6, 8 +c) Prószyński, Deeleman-Reinhold, 2012. Arthr. select. 21(1): 38, 47-58. By courtesy.
Gelotia sp. ©Photo R R. Jackson. By courtesy.
b)
Gelotia argenteolimbata +b) sp [Borneo] : + Proszynski 1987. Atlas ...: 28 [Lectotype] +b) ©Photo P. Koomen. By courtesy.
b)
Gelotia lanka b) robusta : Wijesinghe D.P. 1991b. J. NY Entom. Soc., New York, 99 (2): 274-277, figs 1-6 +b) Wanless 1984a. Bull. brit. Mus. nat. Hist. (Zool.), 46 (2): 174-176, ff. 16C, 19A-F. By courtesy.
b)+
Gelotia salax b) syringopalpis : Wanless 1984a. Bull. brit. Mus. nat. Hist. (Zool.), 46 (2): 176-178, ff. 20A-I +b) 178-180, ff. 21A-I. + Xie, Peng 1995c. Acta zootax. sinica. 20, 3: 289-290, f 1-5. By courtesy.

Meata Żabka, 1985,
known only from the female, is here synonymized with male Gedea Simon, 1902 ,
based on collecting and molecular data
(NEW SYNONYMY) (Maddison, unpublished data).

Gen. Meleon Wanless, 1984 (8 species)

Type species Meleon kenti.
Diagnostic illustrations below are integral part of definition.

Meleon kenti : Wanless F.R. 1978f. Bull. brit. Mus. nat. Hist. (Zool.)., London, 34 (3): 90-91, ff. 2 a-d +34(3): 92, f 3F.
b)
Meleon guinensis +b) insulanus : Azarkina, Logunov 2010. Afr. Invert. 50(1): 171-173, f 38-39 +b) Logunov, Azarkina, 2007: Arthr. Sel. 16(2): 100, f. 17-28. By courtesy.
+b)
Meleon solitaria b) tsaratanana : Wesołowska, Edwards 2012. Ann. zool.: 62(4): 751-752, f 63–65 + Wanless 1978f. Bull. brit. Mus. nat. Hist. (Zool.)., 34 (3): 187-189, f. 26A-G +b) Logunov & Azarkina, 2007[2008b]: 16(2): 103, 36-40. By courtesy.
b)
Meleon madagascarensis +b) raharizonina : Azarkina, Logunov 2010. Afr. Invert. 50(1): 173-176, f 40, 41, 42-47, 53-56 +b) 177, f 37, 48-51. By courtesy.
Meleon russata : Azarkina, Logunov 2010. Afr. Invert. 50(1): 177-180, f 26-27, 57-69. By courtesy.

Gen. Mintonia Wanless, 1984 (9 species)

Type species Mintonia tauricornis.
Diagnostic illustrations below are integral part of definition.

Mintonia sp.: Maddison 2015. Journal of Arachnology. 43: 231–292, f. 11. By courtesy.
+
Mintonia tauricornis : Prószynski, Deeleman-Reinhold, 2012. Arthr. select. 21(1): 43-45, f 87-95
+ Wanless 1984. Bull. brit. Mus. nat. Hist. (Zool.), 46 (2): 158-160, ff. 8A-H, 31A, 32E-F, 35B.. By courtesy.
b)c)
Mintonia breviramus +b) caliginosa +c) mackiei : Wanless F.R. 1984a. Bull. brit. Mus. nat. Hist. (Zool.), 46 (2): 164-165, ff. 12A-E +b) 118-119, f. 8A-E, 19A-C +c) 160-161, ff. 9A-D. By courtesy.
b)c)d)
Mintonia melinauensis +b) nubilis +c) protuberans +d) silvicola :Wanless 1987a. Bull. brit. Mus. nat. Hist. (Zool.). 52 (3): 116, ff. 7A-E +b) 161-162, ff. 10A-F +c) 162-64, ff. 11A-F +d) 119-121, ff. 9A-G. By courtesy.
Mintonia ramipalpis : Wanless F.R. 1984. Bull. brit. Mus. nat. Hist. (Zool.), 46 (2): 166-169, f. 14A-I, 15A-C, 31B-F, 34A-B, 35E-F, 36F. By courtesy.
b)
Mintonia bani +b) sp [Australia] : Ikeda, H. 1995b. Acta. arachn. 44: 117-121, figs 1-6.
+b) Davies Todd, Zabka 1989. Mem. Queensland Mus., 27 (2): 197, t. 5. By courtesy.
b)
Mintonia sp [AUS-1256] +b) sp [Borneo] : Queensland. ©Photo R. Whyte and I.R. Macaulay +b) ©Photo P. Koomen. By courtesy.
Mintonia ignota :Logunov & Azarkina, 2008a: 103, f. 41-51 (Dm). By courtesy.

Gen. Neobrettus Wanless, 1984 (4 species)

Type species Neobrettus tibialis .
Diagnostic illustrations below are integral part of definition.

Neobrettus sp.: Maddison 2015. Journal of Arachnology. 43: 231–292, f. 10. By courtesy.
++
Neobrettus tibialis : Proszynski 1978a. Ent. bas. 19-21, f 25-27 + Wanless 1984a. Bull. brit. Mus. nat. Hist. (Zool.), 46 (2): 183-185, f 24A-H + ©Photo John Murphy, 1997. By courtesy.
b)
Neobrettus phui +b) xanthophyllum : Zabka M. 1985. Ann. zool. 39, 11: 439, f 502-507 +b) Deeleman-Reinhold, Floren. 2003. Bull. British Arachnol. Soc. 12 (7):7. By courtesy.
Neobrettus cornutus : Deeleman-Reinhold, Floren. 2003. Bull. British Arachnol. Soc. 12 (7): 343, f 27-35. By courtesy.
>
Neobrettus sp [Philippines] ©Photo R.R. Jackson [Philippines: Mt. Banaham]. By courtesy.

Gen. Paracyrba Żabka, Kovac 1996 = Cyrba Simon, 1876 (syn. n.)

Paracyrba wanlessi = Cyrba wanlessi (comb. n.) .

Gen. Phaeacius Simon, 1900 (12 species)

Type species Phaeacius fimbriatus.
Diagnostic illustrations below are integral part of definition.

+
Phaeacius fimbriatus : Prószynski, Deeleman-Reinhold, 2012. Arthr. select.21(1): 55, f 130-140 + .19(3): 178, 127-128 + Wanless F.R. 1981b. Bull. brit. Mus. nat. Hist. (Zool.), 41 (4): 202-205, ff. 4A-H, 5B. By courtesy.
+
Phaeacius lancearius : Prószynski, Deeleman-Reinhold, 2012. Arthr. select.21(1): 55, f 130-140 + .19(3): 178, 127-128.+ Wanless F.R. 1981b. Bull. brit. Mus. nat. Hist. (Zool.), 41 (4): 205, f. 1, 2A-F, 5A, C-F. By courtesy.
++
++
Phaeacius malayensis : Zhang J., Li D. 2005. 53: 221-229 + .19(3): 178, 127-128 + Wanless 1981b. 41 (4): 205-208, f 6A-E; 7A-C..+ ©Photo J. Koh 1989 + © Photo H.K. Tang + ©Photo Hirzi Hussain + ©Photo Murphy F. and J. 2000. By courtesy.
b)+
Phaeacius azarkinae +b) alabangensis :Prószynski, Deeleman-Reinhold, 2012. Arthr. select. 19(3): 178, 127-128 +b) Wijesinghe 1991a. Bull. British Arachn. Soc. 8 (8): 252-253, f 9-10 + ©Photo M. Freudenschuess. By courtesy.
b)c)
Phaeacius biramosus +b) leytensis +c) wanlessi : Wijesinghe 1991a. Bull. British Arachn. Soc., 8 (8): 251-252, f 6-8 +b) 253-254, f 11-12 +c) 294-251, f 1-5. By courtesy.
+b)
Phaeacius mainitensis +b) saxicola : Prószynski (Internet) sketch of the type specimen + Barrion A.T. Litsinger J. A. 1995. 45-47, figs 16a-f +b)Wanless 1981b. Bull. brit. Mus. nat. Hist. (Zool.),41 (4): 210, f 9A-C. By courtesy.
b)c)
Phaeacius canalis +b) yixin +c) yunnanensis : Wanless 1981b. Bull. brit. Mus. nat. Hist. (Zool.), 41 (4): 209-210, f 8A-G +b) Zhang J., Li D. 2005. 225, f. 3A-E +c) Peng X., Kim J.P. 1998. Korean J. Biol. Sci., 2: 412, f 2a-d.. By courtesy.

Gen. Portia Karsch, 1878 (14 species)

Type species Portia fimbriata.
Diagnostic illustrations below are integral part of definition.

>
Portia fimbriata : Wanless 1978f. Bull. brit. Mus. nat. Hist. (Zool.). 99-100, f 7A-G; 8A-F; t 3a-f; 4c-f; 5c, d, f + Davies Todd V., Zabka M. 1989. Mem. Queens. 27 (2): 195, t 3 + Photo © Photo Marcus Ng. + H.K. Tang. By courtesy.
Portia labiata : Proszynski 1984c: 26, 111 +Prószynski, Deeleman-Reinhold, 2012. Arthr. select. 21(1): 43-45, f 87-95 + © Photo Koh J. + H.K. Tang. By courtesy.
>
Portia africana : Proszynski 1984c: 99 + Wesołowska, Tomasiewicz, 2008. J. Afrotr. Zool. 4: 45, f 171-173 + © Photo R.R. Jackson. By courtesy.
+
Portia schultzi : Wanless 1978f. Bull. brit. Mus. nat. Hist. (Zool.), London. 88-90, f 1A-B + Wesolowska, Cumming 2008. Ann. zool. 58: 207-208, 131-134.. By courtesy.
++b)
Portia albimana +b) taiwanica : Wanless 1978f. Bull. brit. Mus. nat. Hist. (Zool.). 34 (3): 107-108, f 12A-D + Zabka 1985. Ann. zool., 39, 11: 437, f 487-490 + Murphy, Murphy 1983a. Bull. Br. arachnol. Soc.: 40, f 4-5 +b) Zhang J., Li D., 2005: 53: 226,f. 4A-G. By courtesy.
b)c) d) e)
Portia assamensis +b) crassipalpis +c) hoggi +d) quei +e) jianfeng: Wanless 1978f. Bull. brit. Mus. nat. Hist. 105-107, f 10 d-f, 11 d-f +b) 100-103, f 9A-D; t 5a, e. +c) Zabka 1985. Ann. zool. 39, 11: 437-438, f 491-496 +d) 438, f 497-501 +e) Song, Zhu, Chen 1999: : 541, figs 311K-L . By courtesy.
b)c)
Portia orientalis +b) songi +c) heteroidea: Murphy, Murphy 1983. Bull. Br. arachnol. Soc.: 40, f 6, 9, 12 +b) Song, Zhu, Chen 1999: 541, f 311O-P, 312H, 313A, 328 Q +c) Peng, Xie, Xiao 1993: 187-188, f 653-659. By courtesy.

Gen. Sparbambus Zhang J., Woon J., Li D., 2006 (1 species)

Type species Sparbambus gombakensis. Drawings below are integral part of the definition.

Sparbambus gombakensis Zhang, Woon, Li 2006. Raff. Bull. Zool., 54(2): 241-244, f 1-11.. By courtesy.

Gen. Spartaeus Thorell, 1891 (13 species)

Type species Spartaeus spinimanus.
Diagnostic illustrations below are integral part of definition.


>>
Spartaeus spinimanus : Prószynski, Deeleman-Reinhold, 2010. Arthr. selecta.19(3): 180, f 134-137 + Wanless 1984. Bull. brit. Mus. nat. Hist. (Zool.), 46 (2): 148-151, f 3A-F, 4A-G, 30A-D, 33F, 35A +©Photo D. Petot. By courtesy
b)c)d)
Spartaeus abramovi +b) banthamus +c) jaegeri +d) noctivagus : Logunov Azarkina 2007[2008b]: Arthr. Sel. 16(2): 104-108, f 52-56 +b) 108, f 57-60 +c) 108, f 61-55 +d) 108, f 71-85 ©Photo D. Petot. By courtesy
++
Spartaeus thailandicus : Prószynski Internet drawings: Khao Yai National Park, coll. Deeleman-Reinhold + Logunov, Azarkina 2007: Arthr. Sel. 16(2): 110, f 86-94 + Wanless 1984. Bull. brit. Mus. nat. Hist. (Zool.), 46 (2): 151-152, f 5A-D. By courtesy.
++
Spartaeus uplandicus : Prószynski Internet sketches of holotype female and a congeneric male specimens, coll. IRRI, Los Banos + Barrion A.T., Litsinger . 1995: 43, f 15a-g. By courtesy
b)+
Spartaeus wildtrackii +b) zhangi : Wanless 1987a. Bull. brit. Mus. nat. Hist. (Zool.)., 52 (3): 112-116; f 6A-E. +b) Logunov, Azarkina 2007]: Arthr. Sel. 16(2): 110, f 95-106 + Peng X., Li S. 2002c. Orient. 36: 398-399, f 4a-f. By courtesy
>
Spartaeus platnicki +b) jianfengensis : Peng X., Li S. 2002c. Oriental Insects, 36: 395, f 2a-g +b) 395, f 1a-g. By courtesy
>
Spartaeus emeishan +b) ellipticus : Zhu M., Yang Z., Zhang Z., 2007. J. Hebei Univ. (Nat. Sci. Ed.) 27(5): 515-517, f 2A-H. + Bao Y., Peng X. 2002. Zoological Studies 41 (4): 409-410, f 22-25. By courtesy

Gen. Taraxella Wanless, 1984 (5 species)

Type species Taraxella solitaria.
Diagnostic illustrations below are integral part of definition.

b)>
Taraxella solitaria +b) sumatrana :Wanless, 1987 Bull. British Mus. (NH) (Zool. ser.) 52(3): 156-157, f 7A-G +b) 125-128, t 12 A-J + Prószynski Deeleman-Reinhold, 2012. Arthr. select. 21(1): 57-58, f 154-158. By courtesy.
>
Taraxella hillyardi +b) petrensis :Wanless, 1987 Bull. British Mus. (NH) (Zool. ser.) 52(3): 122-123, f 10A-F. +b) 123-125, f 11A-J. By courtesy.
Taraxella reinholdae :Wanless, 1987 Bull. British Mus. (NH) (Zool. ser.) 52(3): 128, f 13A-G. By courtesy.

Gen. Veissella Wanless, 1984 (2 species)

Type species Veissella durbani.
Diagnostic illustrations below are integral part of definition.

+b)
Veissella durbani +b) milloti :Wesolowska & Haddad, 2009: 95, f. 205-209 + Wanless 1984a. Bull. brit. Mus. nat. Hist. (Zool.), 46 (2): 189, f. 27A-G + Logunov & Azarkina, 2007: 16(2): 113, f 107-113. By courtesy.

Gen. Wanlessia Wijesinghe, 1992 (1 species)

Type species Wanlessia denticulata.
Diagnostic illustrations below are integral part of definition..

Wanlessia denticulata +b)sedgwicki : Peng X., Tso I., Li S. 2002. Zoological Studies 41 (1): 5, f 17-20 +b) Wijesinghe, 1992. Raffles Bulletin of Zoology 40(1): 11, f 1-11. By courtesy.

Gen. Yaginumanis Wanless, 1984 (3 species)

Type species Yaginumanis sexdentatus.
Diagnostic illustrations below are integral part of definition.

>>b)
Yaginumanis sexdentatus +b) saganus [interpreted by Wanless as type of Y. sexdentatus] : Bohdanowicz, Proszynski 1987. Ann. zool. 41, 2: 46-47, f 1-5 +b) Wanless 1984.1987. Bull. brit. Mus. nat. Hist. (Zool.), 46 (2): 153-155, f 6g-i . By courtesy.
b)
Yaginumanis cheni +b) wanlessi : Peng X., Li S. 2002d. Acta zootax. Sinica, 27, 2: 238-240, f 1-4 +b) Zhang J., Li D. 2005: 53: 227, f. 5A-E. By courtesy.