Maddison's alternative

RETURN TO DATABASE - ALTERNATIVE CLASSIFICATION
SALTICIDAE OF THE WORLD - 0-INDEX of Genera - 01-FOREWORD & KEY to groups of genera! - 02-Comparison - HISTORICAL classification of Aelurilleae - 03-AEURILLINES - 04-AMYCINES- 05-AMYCOIDA VARIA - 06-ASTIAINES - BALLINES - 07-BELIPPINES - 08-CHRYSILLINES - 09-COCALODINES - 10-COLONINES[=Thiodinines] - 11-DENDRYPHANTINES - 12-DIOLENINES - EUODENINES - 13-EUOPHRYINES-PART 0: Introduction-Diversity - 13-EUOPHRYINES-PART 1: Admestina-Donoessus - 14-EUOPHRYINES-PART 2: Echeclus-Pystira - 15-EUOPHRYINES-PART 3: Rhyphelia-Zenodorus - 16-EUPOAINES - 17-EVARCHINES - 18-FOSSILS - 19-HABRONATTINES - 20-HARMOCHIRINES - 21-HELIOPHANINES - 22-HISPONINES - 23-HYLLINES - 24-ICIINES - 25-LAPSIINES - 26-LIGONIPEINES - 27-LYSSOMANINES - 28-MENEMERINES - 29-MYRMARACHNINES - 30-NOTICIINES - 31-PELLENINES - 32-PSEUDICIINES - 33-SIMAETHINES - 34-SITTICINES - 35-SPARTAEINES - 35a-SYNAGELINES - 36-THIRATOSCIRTINAE - 37-YAGINUMAELLINES - 38-YLLENINES - 39-UNCLASSIFIED TEMPORARY
APPENDICES -AP1-Simon's classification - AP2-Petrunkevich synthesis-1928 - AP3-Bonnet's list of subfamilies - AP4-Prószynski's revision of subfamilies 1976 - AP5-Prószynski - summary of results - AP6-Aelurillus black & gray - AP7-Maddison's views on Salticidae phylogeny 2014 - MS1-Omoedus synonymy
MADDISON'S ALTERNATIVE: Tittle_pg Index Introduction Agoriini Aelurillina Asemoneinae Amycini Amycoida-incertae sedis Astiini Ballini Baviini Bredini Chrysillini Cocalodini Dendryphantina Euophryini-1 Euophryini-2 Euophryini-3 Eupoinae Freyina Gophoini Harmochirina Hasariini Hisponinae Holcolaetina Huriini Itatina Lapsiini Leptorchestini Lyssomaninae Marpissina Mopsini Myrmarachnini Nannenini Neonini Onomastinae Plexippina Salticini Sarindini Scopocirini Simaethina Simonellini Sitticini Spartaeina Synagelina Thiodinini Thiratoscirtina Tisanibini Viciriini Salticidae-incertae sedis

Monograph of the Salticidae (Araneae) of the World 1995-2015. Part I: Introduction to alternative classification of Salticidae.

Comparison of classifications by Maddison 2015 and Prószyński 2016a
Compiled by Jerzy Prószyński

Maddison's alternative
M_14 - Sitticini
Subfamily: Salticinae: Clade: Amycoida: Tribe: Sitticini
Version July 1st, 2016.
Sources and permissions of illustrations are displayed in the IInd part of the monograph http://www.peckhamia.com/salticidae/

Type genus Sitticus.
Interactive index of genera Aillutticus Amatorculus Attulus Capeta Gavarilla Jollas Nosferattus Pseudattulus Semiopyla Sitticus Sitticus (distinguendus group) Sitticus (floricola group) Sitticus (leucoproctus group) Sitticus (longipes group) Sitticus(palpalis group) Sitticus (saxicola group) Sitticus (terebratus group) Sittipub

Gen. Aillutticus Galiano, 1987 (8 species)

Type species Aillutticus nitens
Diagnostic drawings below are integral part of definition.

Aillutticus sp. : Maddison 2015. Journal of Arachnology. 43:231–292, f. 32. By courtesy.

Aillutticus nitens: Galiano 1987a. Bull. Br. arachn. Soc. 7 (5): 160-162, f 1-2, 6-11, 18, 20, 24. By courtesy.

Aillutticus pinquidor: Galiano 1987a. Bull. Br. arachn. Soc. 7 (5): 160-162, f 1-2, 6-11, 18, 20, 24. By courtesy.

Aillutticus knysakae +b) montanus +c) raizeri : Ruiz, Brescovit 2006c. Revta bras. Zool. 23(2): 530-532, f 3-6 +b) 532, f 7-10 +c) 532-534, f 11-14. By courtesy.

Aillutticus soteropolitano +b) viripotens +c)rotundus : Ruiz, Brescovit 2006c. Revta bras. Zool. 23(2): 534, f 15-18 +b) 534, f 19-22 +b) 530, f 1-2 + Galiano 1987a. Bull. Br. arachn. Soc. 7 (5): 164, ff. 14, 17. By courtesy.

Gen. Amatorculus Ruiz & Brescovit, 2005 (2 species)

Type species Amatorculus stygius.
Diagnostic drawings below are integral part of definition.

Amatorculus stygius + cristinae : Ruiz, Brescovit 2005a. Revta bras. Zool. 22: 694, f 25-28 + 354-355, f 13-16. By courtesy.

Attulus Simon, 1889*

Type species Attulus helveolus Simon 1937: 1198, 1258 = Attus distinguendus Simon 1868: 540 [= Sitticus distinguendus (Simon 1868: 540, Prószyński 1987: 94-95)].
[Comment by Prószyński: synonymy documented in the Atlas 1987, close relationship with 22 species of the Sitticus distinguendus group of species. ]

Gen. Capeta Ruiz & Brescovit, 2005 (2 species)

Type species Capeta tridens.
Diagnostic drawings below are integral part of definition.

Capeta tridens +b) cachimbo : Ruiz , Brescovit. 2005a. Revta bras. Zool. 22: 692, f 21-24 +b) Ruiz, Brescovit 2006a.
Revta bras. Zool. 23(2): 353-354, f 9-12. By courtesy.

Gen. Gavarilla Ruiz, Brescovit 2006 (2 species)

Type species Gavarilla ianuzziae.
Diagnostic drawings below are integral part of definition.

Gavarilla ianuzziae +b) aretada :Ruiz, Brescovit 2006a. Revta bras. Zool. 23(2): 351, f 1-4 +b) 353, f 5-8. By courtesy.

Gen. Jollas Simon, 1901 ( 5 species)

Type species Jollas geniculatus.
Diagnostic drawings below are integral part of definition.

Jollas sp.: Maddison 2015. Journal of Arachnology. 43: 231–292, f. 33. By courtesy.

Jollas geniculatus : Proszynski J. 1987. Atlas ...: fig. 58-59 ( "21085 Jollas geniculatus" "obscurus" - Mus. Paris)
+ Galiano 1991b. Physis C. 47 (112): 18, 19, ff. 1, 5, 14-18, 25, 35, 38, 40-41. By courtesy.

Jollas amazonicus: Galiano 1991b. Physis C. 47 (112): 22, ff. 2, 13, 24, 27. By courtesy.
"Jollas" armatus + crassus : Bryant 1943. Bull. Mus. Comp. Zool. 92 (9): 495, T. 7, F. 70, 73 +497, T. 6, F. 58, 60, 67 . MISPLACED = EUOPHRYINAE.

Jollas manantiales : Galiano 1991b. Physis C. 47 (112): 27, ff. 9-11, 19-20, 28, 39, 46-47. By courtesy.

Jollas pompatus : Proszynski J. 1987. Atlas ...: 58 ("9269 Jollas - Caracas" - Mus. Paris) + Galiano 1991b. Physis C. 47 (112): 18, 19, ff. 1, 5, 14-18, 25, 35, 38, 40-41. By courtesy.

Jollas paranacito + puntalara: Galiano 1991b. Physis C. 47 (112): 23, ff. 42-43 + 23, ff. 4, 6, 21, 29-34, 36. 48-49. By courtesy.

Gen. Nosferattus Ruiz & Brescovit, 2005 (5 species)

Type species Nosferattus discus. Placement uncertain. Diagnostic drawings below are integral part of definition.

Nosferattus discus +b) aegis +c) ciliatus : Ruiz, Brescovit 2005a. Revta bras. Zool. 22(3): 688, f 1-5 +b) 689, f 6-9 +c) 691, f 15-16. By courtesy.

Nosferattus occultus +b) palmatus : Ruiz, Brescovit 2005a. Revta bras. Zool. 22(3): 690, f 10-14 +b)692, f 17-20 . By courtesy.

Gen. Pseudattulus Caporiacco, 1947 (1 species)

Type species Pseudattulus kratochvili.
Diagnostic drawings below are integral part of definition.

Pseudattulus kratochvili +b) [SUPPOSED SYNONYM?] Sitticus cabellensis : Ruiz, Brescovit & Lise, 2007: 377 +b) Proszynski J. 1971a. Annales zoologici, 28: 198-202, f 31-39. By courtesy. NOTE. Ruiz et al. (2005c), propose to synonymize Pseudattulus incertus with female of Sitticus cabellensis (2007) stating bluntly that the holotype specimen of the latter WAS NOT EXAMINED, they do not produce any documentation of females of both species. FOR THESE REASONS THE SYNONIMISATION CANNOT BE ACCEPTED].

Gen. Semiopyla Simon, 1901 ( 3 species)
PLACEMENT?

Type species Semiopyla cataphracta.
Diagnostic drawings below are integral part of definition.

Semiopyla cataphracta :[SUBFAMILY ?] Galiano 1963b. Physis, 23 (66): 442, t. 34, f 8-12 + Galiano 1985c. Rev. suisse
Zool., 92 (2): 282-284, f 1-5, 20 + Cutler 1971b. Ent. News, 82: 141-146, f 1-6. By courtesy.

Semiopyla triarmata : Galiano 1985c. Rev. suisse Zool., 92 (2):287, 290, f 9, 12-13, 17-19. By courtesy.

Semiopyla viperina : Galiano 1985c. Rev. suisse Zool., 92 (2): 284-287, f 6-8,10-11, 14-16, 21-23. By courtesy.

Sitticus Simon, 1901

Type species Sitticus terebratus.

Sitticus (terebratus group = nominal group)

Representative species (and the type species of the genus) Sitticus terebratus [= syn. Araneus terebratus Clerck, 1757].

Sitticus (terebratus group) terebratus : Proszynski J. 1968d. Ann. zool. 26: 396-399, f 1-2, 4-8 [drawing of copulatory ducts simplified] + Proszynski 1983. Verh. naturwiss. Verein Hamburg, (NF) 26; 161-179, map 8 + © Photo J. Lissner. By courtesy.

Sitticus (terebratus group) fasciger :Proszynski J. 1968d. Ann. zool. 26: 399-402, f 3, 9-16. + Proszynski 1983. Verh. naturwiss. Verein Hamburg, (NF) 26; 161-179, map 8. By courtesy. COMPARE Maevia expansa - analogy in palp! (but epigyne unknown) .

Sitticus (terebratus group) finschi : Proszynski 1968d. Ann. zool. 26: 402-406, f 17-23 + Proszynski 1983. Verh. naturwiss. Verein Hamburg, (NF) 26; 161-179, map 8 . By courtesy. COMPARE Maevia expansa - analogy in palp! (but epigyne unknown)!

Sitticus (terebratus group) godlewskii + tannuolana: Proszynski J. 1962a. Bull. Acad. Polon. Sci. 10: 65-68, f 3-4
+ Logunov 1991. Zool. zh., 70 (2): 52, f 4-6. By courtesy.

Sitticus (distinguendus group) (22 species)

Representative species Sitticus distinguendus Simon, 1868 [= Attus distinguendus Simon, 1868].

Sitticus (distinguendus group) distinguendus : Proszynski 1987a. Atlas ...: 94-95 + © Photo J. Lissner. By courtesy.

Sitticus (distinguendus group) ammophilus : Proszynski 1987a. Atlas ...: 94-95 + © Photo J. Lissner. By courtesy.

Sitticus (distinguendus group) ansobicus (+ S. "pubescens" by Caporiacco) + : Proszynski 1987a. Atlas ...: 88-89 + Proszynski 1983. Verh. naturwiss. Verein Hamburg, (NF) 26; 161-179. By courtesy.

Sitticus (distinguendus group) avocator: Proszynski 1987a. Atlas ...: 90-92 (diversity) + Proszynski 1983. Verh. naturwiss. Verein Hamburg, (NF) 26; 161-179. By courtesy.

Sitticus (distinguendus group) albolineatus : Proszynski J. 1987a. Atlas ...: 85-86. + Proszynski J. 1979. Annales zoologici, 34: 316-317, f 286-289 + Proszynski 1983. Verh. naturwiss. Verein Hamburg, (NF) 26: 161-179 + Logunov, Marusik 2000b. KMK Sci. Press Ltd. 202, map 47. By courtesy.

Sitticus (distinguendus group) damini : Proszynski 1987a. Atlas ...: 93. By courtesy.

Sitticus (distinguendus group) niveosignatus : Proszynski 1987a. Atlas ...: 97 + Proszynski 1983. Verh. naturwiss. Verein Hamburg, (NF) 26; 161-179 + Zabka 1980a. Senck. biol., 60 (3/4): 241-247, f 1-19. By courtesy.

Sitticus (distinguendus group) vilis + goricus : Proszynski 1987a. Atlas ...: 98-99. By courtesy.

Sitticus (distinguendus group) burjaticus + dubatolovi+ inopinabilis : Logunov, Danilov 1993. Arthr. Sel., 2 (4): 35-36, f 4a-e. + Logunov, Rakov. 1998. Arthr. Sel., 7(2): 139-140, f 73-77 + Logunov 1992a. Arthr. Sel., 1 (1): 57, f. 6c-d + Logunov, 1998d. Zoosyst. Rossica, 7(1): 78, f 1-2. By courtesy.

Sitticus (distinguendus group) karakumensis + kazakhstanicus + zaisanicus: Logunov 1992a. Arthr. Sel., 1 (1): 59, f 7a-d + 62, f. 8a-d + Logunov, Rakov 1998. Arthr. Sel., 7(2): 140, f. 49-50 + Logunov 1998d. Zoosyst. Rossica, 7(1): 80, f 3-4.

Sitticus (distinguendus group) nenilini +b) eskovi : Logunov, Wesolowska 1993. Ent Basiliensia, 16: 9, f 3-4, 9-12 +b) Logunov , Wesolowska 1995. Genus, 6 (2): 164-167, f 1-6, 8. By courtesy.

Sitticus (distinguendus group) talgarensis + sinensis: Logunov, Wesolowska 1993. Ent Basil., 16: 5-9, f 1-2, 5-8 + Peng X., Xie L., Xiao X. 1993. 44: 219-220, f 778-786. By courtesy.

Sitticus (distinguendus group) penicilloides : Wesolowska 1981a. Annales zoologici, 36: 79-80, f 108-109 + Logunov 1993. Genus, 4 (1): 1-15,f 1-39. By courtesy.

Sitticus penicillatus : Proszynski J. 1973a. Ann. zool. 30: 72-79, f 1-16 + © Photo J. Lissner By courtesy.

Sitticus saltator: Proszynski 1987: 89 + Zabka 1997. F. Polski 19: 94-95, f 358-363 + Proszynski 1983. Verh. naturwiss. Verein Hamburg, (NF) 26; 161-179, map 8 + © Photo J. Lissner + Arachnol. Gesells. By courtesy.

Sitticus clavator + mirandus : Proszynski 1973a. Ann. zool. 30: 72, f 9-10 + Proszynski 1983. Verh. naturw.
Ver. Hamburg, (NF) 26; 161-179, map 15 + Logunov 1993a. Genus, 4 (1): 10, f 2, 12, 34-39. By courtesy.

Sitticus (floricola group)

Representative species Sitticus floricola [= Euophrys floricola Koch C.L., 1837].

Sitticus (floricola group) floricola : Proszynski J. 1980 Annales zoologici, 36: 10-13, f 3-4, 10-14, 20-21+ + Proszynski
1983. Verh. naturwiss. Verein Hamburg, (NF) 26; 161-179. © Photo J. Lissner. By courtesy.

Sitticus (floricola group) floricola palustris: Proszynski J. 1980 Annales zoologici, 36: 12-14, f 15-19, 22-43
+ Kaston B. J. 1948. 70: 458-458, t. 89, f 1660-1662. © Photo J. Lissner. By courtesy
[Uncertain status with S. floricola: conspecific? subspecific? separate species?].

Sitticus (floricola group) rupicola : Proszynski 1980 Ann. zool., 36: 14-18; f 5-6, 9, 44-57, 58-59 [local variation in epigyne?
different species?] + Logunov, Kronestedt 1997. Bull. British Arachn. Soc., 10 (7): 226, f 29. © Photo J. Lissner. By courtesy.

+ sp?

Sitticus (floricola group) inexpectus : Logunov, Kronestedt 1997. Bull. Brit. Arachn. Soc. 10 (7): 226-232, f 1-49 + © Photo J. Lissner + A. Senglet. By courtesy

Sitticus (floricola group) caricis : Proszynski J. 1980 Annales zoologici, 36: 18-20, f 1-2, 7, 60-63. © Photo J. Lissner. By courtesy.

Sitticus (floricola group) caricis : Proszynski J. 1980 Annales zoologici, 36: 18-20, f 1-2, 7, 60-63. By courtesy.

Sitticus (flor. gr.) zimmermanni : Proszynski J. 1980 Ann. zool., 36: 10-13, f 3-4, 10-14, 20-21 + © Photo J. Lissner. By courtesy

Sitticus (floricola group) atricapillus, Proszynski J. 1980 Ann. zool., 36: 10-13, f 3-4, 10-14, 20-21 (synonyms) + Kronestedt, Logunov. 2003. Rev. suisse Zool. 110 (4): 855-837, f 1-22 (comparison). By courtesy[Differs by white ornaments, hair-splitting differences in palps and epigyne - rather variant of S. zimmermanni ?].

Sitticus (floricola group) cutleri : Proszynski J. 1980 Ann. zool., 36: 130, 32, f 94-95, 96-98 + Internet + Proszynski
1983. Verh. naturwiss. Verein Hamburg, (NF) 26; 161-179. By courtesy.

Sitticus(floricola group) dudkoi + monstrabilis : Logunov, 1998d. Zoosyst. Rossica, 7(1): 77-78, f 5-6. + Logunov 1992a. Arthr. Sel., 1 (1): 64, f 9a-e + Logunov, Kronestedt 1997. Bull. Brit. Arachn. Soc. 10 (7): 232, f 50-51 .

Sitticus (floricola group) magnus: Proszynski J. 1980 Ann. zool., 36: 21-23, f. 64-68 + Proszynski 1983. Verh. naturwiss. Verein Hamburg, (NF) 26; 161-179. By courtesy.

Sitticus(floricola group) pulchellus: Logunov 1992a. Arthr. Sel., 1 (1): 64, f 6a-b + Logunov, 1995a. Zoosyst. Rossica, 7(1): 3, 2: 244, f 22-24 (female sp-?). By courtesy.

+b)

Sitticus a) striatus +b)syn. rivalis+c) comment on syn. rivalis : a)Proszynski J. 1980 Ann. zool., , 36: 10-13, f 3-4, 10-14, 20-21
+ b) Logunov 2004a. Bull. British Arachn. Soc., 13 (2): 33-34, f 17-21 +c) comment on separating of S. striatus and rivalis - By courtesy. Comment by Prószyński. After studying Logunov's description and documentation again, I am unable to find convincing characters warranting separation of S. striatus and rivalis. J. Proszyński, 2015. Complaints of Logunov on too transparent part of epigyne mounted in Canada Balsam slide: as in other old slides mounted in the Canada Balsam, this slide require dissolution in Xylol or Toluol, rinsing of epigyne itself, perhaps restaining, and examionation as temporary slide. J.P.

Sitticus (leucoproctus group)

Sitticus (leucoproctus group) Simon, 1876.

Sitticus (leucoproctus group) leucoproctus : Galiano M.E. 1989. Rev. Soc. ent. Argentina, 45 (1-4): 259, f 1-2, 8-9, 12-14, 19, 22 + © Photo Simó M. & Rodríguez M.E (Uruguay:Canelones, Marindia: costa arenosa). By courtesy.

Sitticus (leucoproctus group) cellulanus + flabellatus: Galiano M.E. 1989. Rev. Soc. ent. Argentina, 45 (1-4): 262, f. 3-5,
10-11, 16-17, 20, 23-26 = 261, f. 6-7, 15, 18, 21. By courtesy.

Sitticus (longipes group)

Representative species Sitticus longipes (Canestrini, 1873) [= Attus longipes Canestrini, 1873].

Sitticus (longipes group) longipes : Proszynski 1973a. Ann. zool., 30: 89-91, f 50-55 + Proszynski 1983. Verh. naturwiss. Verein
Hamburg, (NF) 26; 161-179, map 15 + © Photo B. Knoflach. By courtesy.

Sitticus (longipes group) absolutus + sp? : Proszynski 1973a. Ann. zool., 30: 79-86, f 17-19, 22- 44. By courtesy. ATTENTION. REPLACEMENT NAME "dorsatus" (forgotten and dubious) for "absolutus" NOT ACCEPTED!

Sitticus (longipes group) cursor +juniperi : Proszynski 1973a. Ann. zool., 30: 86-89, f 20-21, 45-49 + Gertsch, Riechert 1976. Am. Mus. Novit. 2604: 21, f 37-41. By courtesy.

Sitticus (palpalis group) (2 species)

Sitticus (palpalis group) Simon, 1876.
Diagnostic drawings below are integral part of definition.

Sitticus (palpalis group) palpalis : Galiano M.E. 1991. Acta Zool. Lilloana 40, 1: 61, ff. 1-5, 8, 10-16. By courtesy.

Sitticus tenebricus +b) vanvolsemorum : Galiano M.E., Baert L. 1990. Bull. Inst. royal Sci. naturel. de Belgique, Entomologie, 60: 60: 126-127, f 1, 8, 19, 20, map 7 + Baert , 2011. © Bull. Soc. roy. Belge Ent. 147: 63-67, f 6. +b) Baert , 2011. © Bull. Soc. roy. Belge Ent. 147: 63-67 f 1-5. By courtesy.

Sitticus (palpalis group) mazorcanus +b) canus : Proszynski 1973a. Ann. zool. 30: 91-93, f 56-57 + Galiano 1991. Acta Zool. Lilloana 40, 1: 61, ff. 1-5, 8, 10-16 +b) 40, 1: 67, f 7, 9. By courtesy.

Sitticus (palpalis group) phaleratus : Galiano M.E., Baert L. 1990. Bull. Inst. royal Sci. naturel. de Belgique, Entomologie, 60: 130-131, f 3, 6-7, 12, 14, 17-18, map 7. By courtesy.

Sitticus uber : Galiano M.E., Baert L. 1990. Bull. Inst. royal Sci. naturel. de Belgique, Entomologie, 60: 127-130, f 2, 4-5, 9-11, 13, 15-16, map 7. By courtesy.

Sitticus (saxicola group)

Representative species Sitticus saxicola (Koch C.L., 1848) [= Euophrys saxicola Koch C.L., 1848].

Sitticus (saxicola group) saxicola : Proszynski J. 1968d. Ann. zool. 26: 396-399, f 1-2, 4-8 + © Photo J. Lissner By courtesy.

Sitticus (saxicola group) dzieduszyckii : Proszynski J. 1987. Atlas ...: 96 + Proszynski 1983. Verh. naturwiss. Verein Hamburg, (NF) 26; 161-179, map 10. By courtesy.

Sitticus (saxicola group) ranieri: Proszynski J. 1968d. Ann. zool. 26: 396-399, f 1-2, 4-8 + © Photo J. Lissner. By courtesy.

Gen. Sittipub Prószynski, 2016 gen. n.

Type species Sittipub pubescens [= syn. Aranea pubescens Fabricius, 1775].

Sittipub pubescens :Zabka 1997. Fauna Polski 19: 5-187, f 346-351 + Proszynski 1987: 97 + Kaston 1948. 70: 460, t. 91, f 1692-94 + © Photo J. Lissner.By courtesy.

Sittipub relictarius : Proszynski 1987. Atlas ...: 99-100 (Sitticus sp. from Naltschik - later paratypes)
+Logunov 1998. Zoosyst. Ros.,7(1): 78, f 7-10 + Proszynski 1983. Verh. naturw. Ver. Hamburg, (NF) 26; 161-179, map 10. By courtesy.