RETURN TO DATABASE - ALTERNATIVE CLASSIFICATION
SALTICIDAE OF THE WORLD - Abstract - INDEX of Genera - FOREWORD & KEY to groups of genera! - AEURILLINES - Comparison HISTORICAL classification of Aelurilleae - AMYCINES- AMYCOIDA VARIA - ASTIAINES - BELIPPINES - CHRYSILLINES - COCALODINES - COLONINES [=Thiodinines] - DENDRYPHANTINES - DIOLENINES - EUOPHRYINES-PART 1 Introduction-Donoessus- EUOPHRYINES-PART 2 Echeclus-Pystira - EUOPHRYINES- PART 3 Rhyphelia-Zenodorus - EUPOAINES - EVARCHINES - FOSSILS - HABRONATTINES - HARMOCHIRINES - HELIOPHANINES - HISPONINES - HYLLINES - ICIINES - LAPSIINES - LIGONIPEINES - LYSSOMANINES - MENEMERINES - MYRMARACHNINES - NOTICIINES - PELLENINES - PSEUDICIINES - SIMAETHINES - SITTICINES - SPARTAEINES - THIRATOSCIRTINAE - YAGINUMAELLINES - YLLENINES - UNCLASSIFIED TEMPORARY - Maddison's views on Salticidae phylogeny 2014 - Omoedus synonymy
OUT OF DATE CLASSIFICATORY SYSTEMS -Simon's classification - Petrunkevich synthesis-1928 - Bonnet's list of subfamilies - Prószynski's revision of subfamilies 1976.
Monograph
of the Salticidae (Araneae) of the World 1995-2015.
Introduction to alternative classification of Salticidae
Classification of the Salticidae genera of the World
group of genera
MYRMARACHNINES
by Jerzy
Prószyński
Version October 10th, 2017
Note on illustrations. This introduction to classification
of Salticidae includes only species defined by diagnostic illustrations in
the taxonomic literature, arranged by similarities of embolus and spermophor.
Images are displayed as thumbnails to save space and allow for rapid scanning
of theoretically closely-related species. Thumbnails are small versions
of illustrations in other parts of the monograph, whera are displayed
together with particulars of their sources and permissions.
Gen. Emertonius Peckham & Peckham,
1892 (7
species)
Type
species Emertonius exasperans Peckham & Peckham,1892.
DIAGNOSIS. Correspond with general characteristics of MYRMARACHNINES, which includes differences from several other ant-mimicking groups of Salticidae by exceptional "pipes-like" sclerotized spermathecae, stretching parallel along median axis of epigyne. Copulatory ducts, invisible without staining, make large membranous coils, running from almost indiscernible slit-like copulatory openings, pressed to median septum of epigyne, towards posterior end of spermathecae, near posterior rim of epigyne. Males have oval, or almost circular bulbus, encircled with coils of embolus inside semitranslucent sheath. Spermophor broad, runs along margin of bulbus, making additional loop, small and thin, before entering embolus. Tibial apophysis (RTA) short, twisted corkscrew-like, or straight. Male chelicerae usually disproportionately long, with row of tiny teeth individual row of separate along internal posterior edge, fang is very long. For details
see drawings (below).
More detailed features include
broader and relatively shorter "pipes" of
spermathecae, bow-like bent, terminating anteriorly by globular
swelling (Figs. 97-98), comparable to that in Myrmapana gen.
n. (Fig. 29). Palps resembling Myrma by spermophor having no additional
loop and by straight tibial apophysis, inclined
but not twisted (Fig. 100-101). Type species differs from remaining genera
of MYRMARACHNINES by shape of thorax and by striking color pattern of abdomen
and carapace (Fig. 102), but coloration of other species is unknown.
DESCRIPTION. The cephalic part and anterior thorax in both sexes of the type
species are flat, or almost flat, the posterior slope is
high and step. Abdomen is oval, not constricted. Chelicerae in male are enlarged,
dorsally flattened, stretches horizontally. Tibial apophysis slightly bent
but not twisted. Windows in
epigyne round, relatively small. Pipes of
spermathecae as long as diameter of windows, bow-like bent, pass into almost
round terminal swellings, which, however is not a discrete compartment. ATTENTION: spermathecae of some Toxeus are almost identical with some Emertonius, while their external appearance is strikingly different! Drawings below are integral part of the definition.
REMARKS. Clear original description and delimitation of the genus by Peckham & Peckham
[ 1892:
54, pl. 4, fig. 8] was blurred by Wanless [1978c] who has overlooked
taxonomic significance of internal structure
of epigyne, of body shape and color pattern and reclassified it to the genus Myrmarachne. Opinion of Wanless was repeated by Edwards & Benjamin
[2009: 22] and
Edwards 2013: 4-5] without adding new evidences. The excellent photographs
by D. Knowles of Emertonius,
supported additionally by photographs of damaged specimen from Bali, confirm
characters presented by Peckham & Peckham. Detailed analysis of properties
of Emertonius was given by Prószyński & Deeleman-Reinhold
[2010: 162-164, Figs 164-171].
REFERENCES. Prószynski J., Deeleman-Reinhold C.L. 2010. Vol. 19. N. 3: P. 162-164. Figs 164-167, 169-171.
Wanless F.R. 1978c. Vol. 33. N. 4. P. 235-238. Figs 1-2. 1a-f, 2a-e.
Simon E. 1901-1903. P. 505.
Emertonius
exasperans +b) wanlessi [in press] +c) koomeni sp. n. [in press] [Sabah].
Comparison of drawing of the type by Peckhams 1892, photo from Bali
and epigyne
of lectotype. Prószyński & Deeleman-Reinhold
[2010], ©Photo D. Knowles +b) Wanless
1978b: 33(4): 235-238, f 1a-f, 2a-e [captioned as Myrmarachne exasperans]
+c) ©Photo P.
Koomen [Borneo: Sabah]. By courtesy.
Emertonius
malayanus . Edmunds & Prószyński
[2003]. By courtesy.
Emertonius
shelfordi: Yamasaki, Ahmad 2013.
Zootaxa 3710 (6): 549-551, f 39A–G, 40A–E. © 2013 Magnolia Press.
By courtesy.
Emertonius
kilifi. Wesolowska,
Russel-Smith. 2000.
Tr. Zool., 13 (1): 72-73, figs 188-191 + Wanless 1978b,
33(4): 102-103, figs 63 c-d, f, j, k, 64a,
d, 65 d.
By courtesy.
Emertonius
laurentinus: Wesołowska,
Haddad 2009. Afr. Invert., 50(1): 61, f 111-114 + Wanless 1978b. 33(4): 99-102, figs 63 a-b, e, g, i, 64 b-c, g-h.
By courtesy.
Defense of separation of Emertonius
from Myrmarachne. Controversy on separation of Emertonius from Myrmarachne: external appearance of type species, spermathecae of 4 Emertonius and 2 Myrmarachne. Also manipulations with the Catalog references and muzzling voice of critics.
For explanation of drawings and sources of illustrations - see other entried on this page. XI. 2016. By courtesy. |